Scientific publications
ORCID: 0000-0001-9670-486X
2024
​
M. Gimeno, J. Gimenz, A. Chiaradria, L.S. Davis, P. Seddon, Y. Ropert-Coudert, R. Reisinger, M. Coll, F. Ramírez (2024) Climate and human stressors on global penguin hotspots: current assessments for future conservation. Global Change Biology; in press
​
M. Gimeno, L. Rossell, L. Julià, J. Giménez, C. Sanpera, M. Coll, P. Bustamante, F. Ramírez (2024) Assessing Mercury Contamination in Southern Hemisphere Marine Ecosystems: The Role of Penguins as Effective Bioindicators. Environmental Pollution; 343:123159. DOI: https://doi.org/10.1016/j.envpol.2023.123159
​
M. Coll, J.M. Bellido, M.G, Pennino, et al. (2024) Retrospective analysis of the pelagic ecosystem of the Western Mediterranean Sea: Drivers, changes and effects. Science of the Total Environment; 907:167790. DOI: https://doi.org/10.1016/j.scitotenv.2023.167790
​
2023
​
E. Fernández-Corredor, L. Francotte, I. Martino, F.A. Fernández-Álvarez, S. García-Barcelona, D. Macías, M. Coll, F. Ramírez, J. Navarro, J. Giménez (2023) Assessing juvenile swordfish (Xiphias gladius) diet as an indicator of marine ecosystem changes in the northwestern Mediterranean Sea. Marine Environmental Research; 192:106190. DOI: https://doi.org/10.1016/j.marenvres.2023.106190
​
F. Ramírez, Y. Cordón, D. García, A. Rodríguez, M. Coll, L.S. Davis, A. Chiaradia, J.L. Carrasco (2023) Large-scale human celebrations increase global light pollution. People and Nature; 00:1–9.. DOI: https://doi.org/10.1002/pan3.10520
​
V. Sbragaglia, L. Espasandín, I. Jaric, R. Vardi, F. Ramírez, M. Coll (2023) Tracking ongoing transboundary marine distributional range shifts in the digital era. Mar. Ecol. Prog. Ser.: SHIFTav3. DOI: https://doi.org/10.3354/meps14309
​
C. Sauser, F. Angelier, F. Ramírez, P. Blévin, O. Chastel, G.W. Gabrielsen, W. Jouanneau, A. Kato, B. Moe, F. Ramírez, S. Tartu, S. Descamps (2023) Demographic responses of Arctic seabirds to spring sea-ice variations. Front. Ecol. Evol., 11: 1107992. DOI: https://doi.org/10.3389/fevo.2023.1107992
​
A. Antoniou, T. Manousaki, F. Ramírez, A. Cariani, R. Cannas, P. Kasapidis, A. Magoulas, M. Albo-Puigserver, E. Lloret-Lloret, J.M. Bellido, M.G. Pennino, M.C. Follesa, A. Esteban, C. Saraux, M. Sbrana, M.T. Spedicato, M. Coll, C.S. Tsigenopoulos (2023) Sardines at a junction: Seascape genomics reveals ecological and oceanographic drivers of variation in the NW Mediterranean Sea. Molecular Ecology, 00: 1-21. DOI: https://doi.org/10.1111/mec.16840
​
2022
​
J. Ouled-Cheikh, F. Ramírez, M. Sánchez-Fortún, A. Cortejna, C. Sanpera, J.L. Carrasco (2022) Fishing activities shape the flight behaviour of an opportunistic predator species. Estuarine, Coastal and Shelf Science, 278:108089. DOI: https://doi.org/10.1016/j.ecss.2022.108089
​
F. Ramírez, L.J. Shannon, C.D. van der Lingen, L. Julià, J. Steenbeek, M. Coll (2022) Climate and fishing simultaneously impact small pelagic fish in the oceans around the southernmost tip of Africa. Frontiers in Marine Science, 9:1031784. DOI: 10.3389/fmars.2022.1031784
​
J. Ouled-Cheikh, M. Coll, L. Cardona, J. Steenbeek, F. Ramírez (2022) Fisheries-enhanced pressure on Mediterranean regions and pelagic species already impacted by climate change. Elementa: Science of the Anthropocene, 10 (1): 00028. DOI: https://doi.org/10.1525/elementa.2022.00028
​
F. Ramírez, L.J. Shannon, R. Angelini, J. Steenbeek, M. Coll (2022) Overfishing species on the move may burden seafood provision in the low-latitude Atlantic Ocean. Science of the Total Environment, 756: 144002. DOI: https://doi.org/10.1016/j.scitotenv.2022.155480
​
F. Ramírez, V. Sbragaglia, K. Soacha, M. Coll, J. Piera (2022) Challenges for Marine Ecological Assessments: Completeness of Findable, Accessible, Interoperable, and Reusable Biodiversity Data in European Seas. Frontiers in Marine Science, 8: 802235. DOI: https://doi.org/10.3389/fmars.2021.802235
​
2021
​
S. Descamps, F. Ramírez (2021) Species and spatial variation in the effects of sea-ice on Arctic seabird populations. Diversity and Distributions, 27: 2204-2217. DOI: https://doi.org/10.1111/ddi.13389
​
J. Ouled-Cheikh, V. Morera-Pujol, Á. Bahillo, F. Ramírez, M. Cerdà-Cuéllar, R. Ramos (2021) Foraging in the Anthropocene: Feeding plasticity of an opportunistic predator revealed by long term monitoring. Ecological Indicators, 129: 107943. DOI: https://doi.org/10.1016/j.ecolind.2021.107943
​
F. Ramírez, A. Chiaradia, D.A. O'Leary, R.D. Reina (2021) Making most of the old age: Autumn breeding as an extra reproductive investment in older seabirds. Ecology and Evolution, 11(10): 5393-5401. DOI: https://doi.org/10.1002/ece3.7431
​
J. Giménez, A. Cañadas, R. de Stephanis, F. Ramírez (2021) Expanding protected areas to encompass the conservation of the endangered common dolphin (Delphinus delphis) in the Alboran Sea. Marine Environmental Reseach, 168: 105305. DOI: https://doi.org/10.1016/j.marenvres.2021.105305.
I. Afán, J.M. Arcos, F. Ramírez, D. García, B. Rodríguez, K. Delord, A. Boué, T. Micol, H. Weimerskirch, M. Louzao (2021) Where to head: environmental conditions shape foraging destinations in a critically endangered seabird. Marine Biology, 168, 23. DOI: https://doi.org/10.1007/s00227-021-03830-1
​
F. Ramírez, D. Vicente-Sastre, I. Afán, J.M. Igual, D. Oro, M.G. Forero (2021) Stable isotopes in seabirds reflect changes in marine productivity patterns. Marine Ecology Progress Series, 662: 169-180. DOI: https://doi.org/10.3354/meps13615
​
I. Paredes, F. Ramírez, D. Aragonés, M.A. Bravo, M.G. Forero, A.J. Green (2021) Ongoing anthropogenic eutrophication of the catchment area threatens the Doñana World Heritage Site (South-west Spain). Wedlands Ecology and Management, 29(1): 41-65. DOI 10.1007/s11273-020-09766-5.
​
​​F. Ramírez, M.G. Pennino, M. Albo-Puigserver, J. Steenbeeck, J.M. Bellido, M. Coll (2021) SOS small pelagics: a Safe Operating Space for small pelagic fish in the Western Mediterranean Sea. Science of the Total Environment, 720: 144002. DOI: //doi.org/10.1016/j.scitotenv.2020.144002.
​
2020
​
J. Ouled-Cheikh, C. Sanpera, J. Bécares, J.M. Arcos, J.L. Carrasco, F. Ramírez (2020) Spatiotemporal analyses on tracking data reveal fine-scale, daily cycles in seabird-fisheries interactions. ICES Journal of Marine Science, fsaa098. DOI: https://doi.org/10.1093/icesjms/fsaa098
​
X. Meyer, A.J.J. MacIntosh, A. Chiaradi, A. Kato, F. Ramírez, C. Sueur, Y. Ropert-Coudert (2020) Oceanic thermal structure mediates dive sequences in a foraging seabird. Ecology and Evolution, 10(13): 6610-6622. DOI: https://doi.org/10.1002/ece3.6393
​
Y. Parra-Torres, F. Ramírez, I. Afán, J. Aguzzi, W. Bouten, M.G. Forero, J. Navarro (2020) Behavioral rhythms of an opportunistic predator living in anthropogenic landscapes. Movement Ecology, 8: 17. DOI: https://doi.org/10.1186/s40462-020-00205-x
​
F. Ramírez, I. Afán, W. Bouten, J.L. Carrasco, M.G. Forero, J. Navarro (2020) Humans shape the year-round distribution and habitat use of an opportunistic scavenger. Ecology and Evolution, 10(11): 4716-4725. DOI: https://doi.org/10.1002/ece3.6226
​
2019
​
P. Lopezosa, M. G. Forero, F. Ramírez, J. Navarro (2019) Individuals within populations: no evidences of individual specialization in the trophic habits of an opportunistic predator. Estuarine, Coastal and Shelf Science, 229: 106427
​
S. Descamps, F. Ramírez, S. Benjaminsen, T. Anker-Nilssen, R.T Barrett, Z. Burr, S. Christensen-Dalsgaard, K.E. Erikstad, D.B. Irons, S.-H. Lorentsen, M.L. Mallory, G. Robertson, H. Strøm, Ø. Varpe and S. Lavergne (2019) Diverging phenological responses of Arctic seabirds to an earlier spring. Global Change Biology, 25(12): 4081-4091. DOI: 10.1111/gcb.14780
​
A. Rodríguez, J.M. Arcos, V. Bretagnolle, M. Dias, N. Holmes, M. Louzao, J. Provencher, A. Raine, F. Ramírez, B. Rodríguez, R.A. Ronconi, R.S. Taylor, E. Bonneaud, S. Borrelle, V. Cortés, S. Descamps, V.L. Friesen, M.Genovart, A. Hedd, P. Hodum, G. Humphries, M. Le Corre, C. Lebarbenchon, R.Martin, E.F. Melvin, W.A. Montevecchi, P. Pinet, I. Pollet, R. Ramos, J. Russell, P.G. Ryan, A. Sanz-Aguilar, D. Spatz, M. Travers, S.C. Votier, R. Wanless, E. Woehler, A Chiaradia (2019) Future directions in conservation research on petrels and shearwaters. Frontiers in Marine Science, 6: 94. DOI:10.3389/fmars.2019.00094.
​
Y. Ropert-Coudert, A. Chiaradia, D.G. Ainley, A. Barbosa, T. Boulinier, R. Brasso, M. Dewar, U. Ellenberg, L. Emmerson, R. Hickcox, S. Jenouvrier, R. McIntosh, L. Phoebe, F. Ramírez, V. Ruoppolo, P.G. Ryan, P. Seddon, R.B. Sherley, R. Vanstreels, L. Waller, E. Woehler, P.N. Trathan (2019) Happy Feet in a hostile world? The future of penguins depends on proactive management of current and predictable threats. Frontiers in Marine Science, 6: 248. DOI:10.3389/fmars.2019.00248
​
J. Hovinen, A. Tarroux, F. Ramírez, M.G. Forero, S. Descamps (2019) Relationships between isotopic ratios, body condition and breeding success in a High Arctic seabird community. Marine Ecology Progress Series, 613:183-195. DOI:https://doi.org/10.3354/meps12886
​
I. Paredes, F. Ramírez, M.G. Forero, A.J. Green (2019) Stable isotopes in helophytes reflect anthropogenic nitrogen pollution in entry streams at the Doñana wetland World Heritage Site. Ecological Indicators, 97: 130-140. DOI:https://doi.org/10.1016/j.ecolind.2018.10.009
​
2018
A. Rodriguez, F. Ramírez, M.N. Carrasco, A. Chiaradia (2018) Seabird plastic ingestion differs among collection methods: examples from the short-tailed shearwater. Environmental Pollution, 243: 1750-1757.
F. Ramírez, M. Coll, J. Navarro, J. Bustamante, A.J. Green (2018) Spatial congruence between multiple stressors in the Mediterranean Sea may reduce its resilience to climate impacts. Scientific Reports, 7: 4500 DOI:10.1038/s41598-017-04775-6
​
J. Giménez, A. Cañadas, F. Ramírez, I. Afán, S. García-Tiscar, C. Fernández-Maldonado, J. José Castillo, R. de Stephanis (2018) Living apart together: niche partitioning among Alboran Sea cetaceans. Ecological Indicators, 95: 32-40
F. Ramírez, C. Rodríguez, J. Seoane, J. Figuerola, J. Bustamante (2018) How will climate change affect endangered Mediterranean waterbirds? PLoS ONE 13(2): e0192702
I. Afán, J. Giménez, MG. Forero, F. Ramírez (2018) An integrated method for identifying marine areas of highest conservation priority. Conservation Biology 0(0): 1-12
2017
Agnew, P., Cannell, B., Carlile, N., Carroll, G., Chiaradia, A., Colombelli-Négrel, D., Copley, P., Dann, P., Grosser, S., Holmberg, R., Houston, D., O’Neill, L., Ramirez, F., Sutherland, D., Tennyson, A., Waugh, S., Wilson, K. & Woehler, E. (2017) Update on little penguin IUCN Red List Assessment. The IUCN Red List of Threatened Species: http://dx.doi.org/10.2305/IUCN.UK.2017-1.RLTS.T22697805A112478911.en
J. Giménez, A. Marçalo, F. Ramírez, P. Verborgh, P. Gauffier, R. Esteban, L. Nicolau, E. González-Ortegón, F. Baldó, C. Vilas, J. Vingada. M.G. Forero, R. Stephanis (2017) Diet of bottlenose dolphins (Tursiops truncatus) from the Gulf of Cadiz: Insights from stomach content and stable isotope analyses. PLoSONE, 12(9): e0184673
J. Giménez, M. Louis, E. Barón, F. Ramírez, P. Verborgh, P. Gauffier, R. Estaban, E. Eljarrat, D. Barceló, M. Forero, R. de Stephanies, (2017) Towards the identification of ecological management units: a multidisciplinary approach for the effective management of bottlenose dolphins in the southern Iberian Peninsula. Aquatic Conservation: Marine and Freshwater Ecosystems doi: 10.1002/aqc.2814
J. Giménez, F. Ramírez, J. Navarro, J. Almunia, M.G. Forero, R. de Stephanis (2017) Lipid effect on isotopic values in bottlenose dolphins (Tursiops truncatus) and its prey with implications for diet assessment. Marine Biology, 164:122
F. Ramírez, A. Tarroux, J. Hovinen, J. Navarro, I. Afán, M.G. Forero, S. Descamps, (2017) Sea ice phenology and primary productivity pulses shape breeding success in Arctic seabirds. Scientific Reports, 7: 4500
F. Ramírez, I. Afán, L. Davis, A. Chiaradia (2017) Climate impacts on global hot spots of marine biodiversity. Science Advances, 3: e1601198
J. Navarro, D. Grémillet, F. Ramírez, I. Afán, W. Bouten, M.G. Forero (2017) Shifting individual habitat specialization of a successful predator living in anthropogenic landscapes. Marine Ecology Progress Series doi: 10.3354/meps12124.
J. Giménez, A. Cañadas, F. Ramírez, I. Afán, S. García-Tiscar, C. Fernández-Maldonado, J.J. Castillo, R. de Stephanis (2017) Intra- and interspecific niche partitioning between sympatric, ecologically similar species. Marine Ecology Progress Series, 567:199-210.
2016
J. Navarro, D. Grémillet, I. Afán, F. Ramírez, W. Bouten, M.G. Forero (2016) Feathered detectives: real-time GPS tracking of scavenging gulls pinpoints illegal waste dumps. PLoSONE, 11(7): e0159974
F. Ramírez, I. Afán, G. Tavecchia, I A. Catalán, D. Oro, A. Sanz-Aguilar (2016) Oceanographic drivers and mistiming processes shape breeding success in a seabird. Proceedings of the Royal Society B, 283: 20152287
C. Coccia, B. Fry, F. Ramírez, L. Boyero, S.E. Bunn, C. Diaz-Salgado, M. Walton, L. Le Vay and A. J. Green (2016) Niche partitioning between invasive and native corixids (Hemiptera, Corixidae) in south-west Spain. Aquatic Sciences, 1: 1-13
A. Chiaradia, F. Ramírez, M.G. Forero, K.A. Hobson (2016) Stable isotopes (δ13C, δ15N) combined with conventional dietary approaches reveal plasticity in central-place foraging behaviour of little penguins Eudyptula minor. Frontiers in Ecology and Evolution 3:154. doi:10.3389/fevo.2015.00154
J. Giménez, F. Ramírez, J. Almunia, M.G. Forero, R. de Stephanis (2016) From the pool to the sea: applicable isotope turnover rates and diet to skin discrimination factors for bottlenose dolphins (Tursiops truncatus). Journal of Experimental Marine Biology and Ecology, 475: 54-61
2015
F. Ramírez, C. Gutiérrez-Expósito, I. Afán, J. Giménez, R. de Stephanis, M.G. Forero (2015) The human influence on gulls’ non-breeding distribution: Potential consequences of changes in fishing practices. Marine Ecology Progress Series 527:221-232.
I. Afán, A. Chiaradia, M.G. Forero, P. Dann, F. Ramírez (2015) A novel spatio-temporal scale based on ocean currents unravel environmental drivers of reproductive timing in a marine predator. Proceedings of the Royal Society B, 282: 20150721.
C. Gutiérrez-Expósito, F. Ramírez, I. Afán, M.G. Forero, K.A. Hobson (2015) Toward a deuterium feather isoscape for sub-Saharan Africa: Progress, challenges and the path ahead. PLoSONE, 10(9): e0135938
F. Ramírez, M.G. Forero, K.A. Hobson, A. Chiaradia (2015) Older female little penguins adjust nutrient allocations to both eggs and chicks. Journal of Experimental Marine Biology and Ecology, 468: 91-96
2014
A. Chiaradia, M.G. Forero, J. McInnes, F. Ramírez (2014) Searching for the true diet of marine predators: Incorporating Bayesian priors into stable isotope mixing models. PLoSONE 9(3): e92665
A. Sanz-Aguilar, G. Tavecchia, I. Afán, F. Ramírez, A. Doxa, A Bertolero, C. Gutiérrez-Expósito, M.G. Forero, D. Oro (2014) Living on the edge: demography of the Slender-billed Gull in the western Mediterranean. PLoSONE 9(3): e92674
​
F. Ramírez, I. Afán, K.A. Hobson, M. Bertellotti, G. Blanco, M.G. Forero (2014) Natural and anthropogenic factors affecting the feeding ecology of a top marine predator, the Magellanic penguin. Ecosphere 5:art38
I. Afán, J. Navarro, L. Cardador, F. Ramírez, A. Kato, B. Rodríguez, Y. Ropert-Coudert, M.G. Forero (2014) Foraging movements and habitat niche of two closely-related seabirds breeding in sympatry. Marine Biology. DOI 10.1007/s00227-013-2368-4
A. Abdennadher, F. Ramírez, M.S. Romdhane, L. Jover, C. Sanpera (2014) Using a three-isotope bayesian mixing model to assess the contribution of refuse dumps in the diet of yellow-legged gull Larus michahellis. Ardeola 62 (1): 297-309
2013
F. Ramírez, M. Garcia-Tarrasón, L. Rami, M. Genovart, L. Jover, C. Sanpera (2013) The role of calcium in constraining egg synthesis in the Audouin's Gull Larus audouinii. Canadian Journal of Zoology 91(9): 640-645
L. Gangoso, R. Marquez-Ferrando, F. Ramírez, I. Gomez-Mestre, J. Figuerola (2013) Understanding phenotypic responses to global change. Report from the Meeting ‘‘Global change in the Mediterranean: learning from experiences worldwide. BioEssays 35(5): 491-495
R. Ramos, F. Ramírez, L. Jover (2013) Trophodynamics of inorganic pollutants in a wide-range feeder, the Yellow-legged gull (Larus michahellis): an isotopic approach for environmental health biomonitoring. Environmental Pollution 172: 235-242
2012
F. Ramírez, J. Navarro, I. Afán, A. Delgado, K.A. Hobson, M.G. Forero (2012) Adapting to a changing world: Unraveling the role of man-made habitats as alternative feeding areas for Slender-billed Gull (Chroicocephalus genei). PLoSONE 7(10): e47551
2011
F. Ramírez, A. Abdennadher, C. Sanpera, L. Jover, L.I. Wassenaar, K.A. Hobson (2011) Assessing waterbird habitat use in coastal evaporative systems using stable isotopes (δ13C, δ15N and δD) as environmental tracers. Estuarine, Coastal and Shelf Science 92(2): 217-222
R. Ramos, F. Ramírez, J.L. Carrasco, L. Jover (2011) Insights into the spatiotemporal component of feeding ecology: an isotopic approach for conservation management sciences. Diversity and Distributions 17(2): 338-349
F. Ramírez, R. Ramos, J.L. Carrasco, C. Sanpera, L. Jover, X. Ruiz (2011) Intra-clutch pattern of albumen δ13C and δ15N in Yellow-legged Gulls Larus michahellis: female dietary shift or resource allocation strategy? Journal of Avian Biology 42(3): 239-246
A. Abdennadher, F. Ramírez, M.S. Romdhane, X. Ruiz, L. Jover, C. Sanpera (2011) Little Egret (Egretta garzetta) as a bioindicator of trace element pollution in Tunisian aquatic ecosystems. Environmental Monitoring and Assessment 175(1): 677-684
F. Ramírez, A. Abdennadher, C. Sanpera, L. Jover, K.A. Hobson, L.I. Wassenaar (2011) Tracing waterbird exposure to trace metals: a case study at the solar saltworks of Thyna (Sfax, Tunisia). Environmental Science & Technology 45(12): 5118-5124
2010
F. Ramírez, K.A. Hobson, O.S. Wangensteen, M. Genovart, G. Viscor, C. Sanpera, L. Jover (2010) A physiological marker for quantifying differential reproductive investment between the sexes in Yellow-legged gulls (Larus michahellis). Journal of Experimental Marine Biology and Ecology 396(1): 48-52
A. Abdennadher, F. Ramírez, M.S. Romdhane, X. Ruiz, L. Jover, C. Sanpera (2010) Biomonitoring of coastal areas in Tunisia: Stable isotope and trace element analysis in the Yellow-legged Gull. Marine Pollution Bulletin 60(3): 440-447
R. Ramos, M. Cerdà-Cuéllar, F. Ramírez, L. Jover, X. Ruiz (2010) The influence of refuse tips exploitation in the prevalence of Campylobacter spp. and Salmonella serovars in seagulls. Applied and Environmental Microbiology 76(9): 3052-3056
2009
F. Ramírez, L. Jover, C. Sanpera, X. Ruiz, E. Piqué, R. Guitart (2009) Combined measurements of egg fatty acids and stable isotopes as indicators of feeding ecology in Lake-dwelling birds. Freshwater Biology 54(9): 1832-1842
R. Ramos, F. Ramírez, C. Sanpera, L. Jover, X. Ruiz (2009) Diet of Yellow-legged Gull (Larus michahellis) chicks along the Spanish Western Mediterranean coast: the relevance of refuse dumps. Journal of Ornithology 150(1): 265-272
R. Ramos, F. Ramírez, C. Sanpera, L. Jover, X. Ruiz (2009) Feeding ecology of yellow-legged gulls Larus michahellis in the Western Mediterranean: A comparative assessment using conventional and isotopic methods. Marine Ecology Progress Series 377: 289-297